Evolutionary Biology

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  • Abitur, Humanistisches Gymnasium Hassfurt, 1969
  • Diplom (M. S.), Physics (minor: Mathematics), University of Erlangen/Nürnberg, 1977
  • Dr. rer. nat. (Ph. D.), Theoretical Biophysics, University of Konstanz, 1981

Honors and Awards

  • DFG Postdoctoral Fellow, 1984
  • NIH Visiting Associate, 1987-89
  • NIH Shannon Award, 1991
  • University of Maryland, Semester Research Award, 1994
  • University of Barcelona, Distinguished Visiting Research Professor, 1996
  • CNRS, Visiting Scientist, Gif-sur-Yvette, 1998
  • Bernhard Rensch Lecture, University of Münster, 2006
  • DFG Research Unit on Natural Selection in Structured Populations, Speaker, 2008-2016
  • DFG Senate Committee on Biodiversity, 2008-16
  • Center for Advanced Studies (CASLMU), LMU Munich, 2011-
  • European Research Council (ERC), Panel Member, 2016-

Academic and Research Appointments

  • Postdoctoral Fellow, Universities of Konstanz and Bremen, 1981-84, and Universities of Sussex and Edinburgh (with J. Maynard Smith and B. Charlesworth), 1984-85
  • Wissenschaftlicher Assistent, University of Technology Darmstadt, 1985-87
  • Visiting Associate, NIH, Research Triangle Park (with C.H. Langley), 1987-89
  • Assistant Professor, Department of Zoology, University of Maryland, College Park, 1989-95
  • Associate Professor, Department of Zoology, University of Maryland, College Park, 1995-97, and Department of Biology, University of Rochester, 1998-99
  • Full Professor, Department of Biology, University of Rochester, 1999-2000
  • Professor and Chair, Department of Biology, University of Munich, 2000-2015
  • Visiting Researcher, Museum für Naturkunde Berlin, 2015-

Extramural Funding

  • NIH Grant, Potential mechanisms of epistatic fitness interactions, 1991-94
  • NSF Grant, Effects of a recent population size bottleneck on genetic variation of whooping cranes (Dissertation Research, Travis Glenn), 1994-96
  • NSF Grant, Effects of natural selection on genetic differentiation, 1994-98
  • NSF Research Training Grant, Biology of small populations (University of Maryland), 1996-2001
  • NIH Grant (with S. Tanda, Ohio University), Evolution by mutations with compensatory fitness interactions, 1998-2002
  • DFG Grant, Die Rolle der natürlichen Selektion in der genetischen Differenzierung von Populationen, 2001-06
  • DFG Grant, A population genetic approach to speciation in the genus Lycopersicon, Schwerpunktprogramm "Radiationen - Genese biologischer Diversität", 2002-08
  • DFG Grant (with G. Heubl, LMU Munich), Origin, evolution and adaptive radiation in the genus Micromeria Benth. (Lamiaceae) in Macaronesia, Schwerpunktprogramm "Radiationen - Genese biologischer Diversität", 2002-04
  • DFG Grant, Nachweis der natürlichen Selektion auf dem X-Chromosom von Drosophila melanogaster, 2002-05
  • VolkswagenStiftung Grant (with D. Tautz, U. of Cologne), Detecting adaptive trait loci via selective sweeps, 2003-06
  • DFG Grant (with L. Rose, LMU Munich), Evolution of a pathogen resistance pathway in the tomato genus, 2005-07
  • DFG Grant, Genetics of adaptation: selective sweeps in Drosophila melanogaster, 2007-10
  • DFG Grant, Population genetics of adaptation in wild tomatoes, 2007-10
  • DFG Grant (with U. Gerland, LMU Munich), Adaptation sensu Sewall Wright: evolution over rugged fitness landscapes, 2007-10
  • DFG Research Unit Grant, Selective sweeps in Drosophila melanogaster, 2008-14
  • DFG Grant (with S. Hutter, LMU Munich), Recent selection on immunity genes in Drosophila, Schwerpunktprogramm "Host-parasite coevolution", 2009-12
  • DFG Grant, Species demography and the distribution of fitness effects of new deleterious mutations in wild tomatoes (Solanum section Lycopersicon, 2010-13
  • DFG Grant, Adaptation genomics of wild tomatoes, 2012-15
  • DFG Grant (with S. Hutter, LMU Munich), Evidence for balancing selection at immunity genes in Drosophila, Schwerpunktprogramm "Host-parasite coevolution", 2012-15
  • DFG Grant, Recurrent selective sweeps versus trench warfare in host-parasite coevolution, Schwerpunktprogramm "Probabilistic Structures in Evolution", 2012-15
  • EU Grant, Marie Curie ITN "Intercrossing", Selective sweep data and next-generation-sequence data analysis, 2012-15
  • Center for Advanced Studies CASLMU (with Gisela Grupe, LMU München), Schwerpunktprogramm "Evolutionary Biology", 2014-16
  • DFG Priority Program (with Karl Schmid, Klaus Fischer, Eva Stukenbrock, and Ralph Tiedemann), "Rapid evolutionary adaptation: Potential and constraints", 2015-21
  • DFG Grant, Modeling and inference of genomic signatures of polygenic selection driving fast adaptation, Schwerpunktprogramm "Rapid evolutionary adaptation: Potential and constraints", 2015-18
  • DFG Grant, The role of recombination in rapid adaptive evolution of fungal plant pathogens: a comparative population genomics study (with PI Eva Stukenbrock, MPI für Evolutionsbiologie, Plön), Schwerpunktprogramm "Rapid evolutionary adaptation: Potential and constraints", 2015-18
  • DFG Grant, Recurrent selective sweeps versus trench warfare in host-parasite coevolution: the influence of population size changes (with Aurélien Tellier, TU Munich), Schwerpunktprogramm "Probabilistic Structures in Evolution", 2016-2018

Editorial Boards

  • Associate Editor, Evolution, 1993-95
  • Associate Editor, Molecular Biology and Evolution, 1998-2003
  • Associate Editor, Genetics, 1998-2003 and 2011-


Ph.D. Students

T. Wiehe (Dr. rer. nat., 1995; DAAD fellow;  now Professor, U. of Cologne)

D. A. Kirby (Ph.D., 1996; now Senior Lecturer, U. of Manchester)

T. C. Glenn (Ph.D., 1997; now Research Scientist, U. of Georgia, SREL)

J. Parsch (Ph.D., 1998; now Professor, LMU Munich)

Y. Kim (Ph.D., 2002; now Associate Professor, Ewha Woman's U., Seoul)

Y. Chen (Ph.D., 2003; Walter-Fitch-Prize 2002; now private sector)

J. F. Baines (Dr. rer. nat., 2004; now Professor, U. of Kiel)

S. Glinka (Dr. rer. nat., 2005; now private sector)

L. Ometto (Dr. rer. nat., 2006; Canestrini Award 2005; now Postdoc, Fondazione Edmund Mach, Italy)

S. Beisswanger (Dr. rer. nat., 2006; now private sector)

U. Arunyawat (Dr. rer. nat., 2007; DAAD fellow; now Lecturer, Kasetsart U., Bangkok, Thailand)

S. Hutter (Dr. rer. nat., 2007; now Research Associate, LMU Munich)

N. Svetec (Dr. rer. nat., 2009; now Postdoc, UC Davis)

L. Grzeskowiak (Dr. rer. nat., 2009; now Postdoc, Fondazione E. Mach, San Michele all'Adige)

C. Merino (Dr. rer. nat., 2009; now Postdoc, U. Peruana Cayetano Heredia)

H. Xia (Ph.D., 2010; Northwest A&F U., China)

A. Hörger  (Dr. rer. nat., 2011; now Assistant Professor, U. of Salzburg)

S. Laurent (Dr. rer. nat., 2011; now Postdoc, EPFL Lausanne)

P. Pavlidis (Dr. rer. nat., Fellow of VW-Foundation, 2011; now Researcher tenure track, Institute of Molecular Biology and Biotechnology (IMBB), Greece)

R. Piskol (Dr. rer. nat., 2011; now private sector)

A. Werzner (Dr. rer. nat., 2011; now DKMS German Bone Marrow Donor Center, Tübingen, Germany)

I. Fischer (Dr. rer. nat., Fellow of Bavarian Eliteförderung, 2012; now Postdoc, SupAgro, Montpellier)

F. Paparazzo (Dr. rer. nat., 2013; now High School Teacher in Milano)

P. Duchén (Dr. rer. nat., 2014; now Postdoc, University of Fribourg)

R. Wilches (Dr. rer. nat., Fellow of VW-Foundation, 2014; now private sector)

S. Voigt (Dr. rer. nat., 2015; now Postdoc, TU Dresden)

K. Böndel (Dr. rer. nat., 2014; now Postdoc, U. of Edinburgh)

V. Božičević (Dr. rer. nat., current)

K. von Heckel (Dr. rer. nat., current)

M. Croze  (Dr. rer. nat., current)



J. M. Braverman (1997-98; Sloan fellow; now Jesuit Brother)

C. Kerdelhué (1998-99; now Chargée de recherche, INRA, Orleans)

D. Carlini (1999-2001; now Associate Professor, American U., Washington, DC)

H. Innan (1999-2000; Japan Society for the Promotion of Science fellow; now Associate Professor, Graduate University for Advanced Studies, Hayama, Japan)

E. Baudry (1999-2000; now Maitre de Conference, U. of Paris)

A. Das (2000-04; now Assistant Director, National Institute of Malaria Research, New Delhi, India)

S. Mohanty (2000-04; now Senior Lecturer, Jaypee Institute of Information Technology U., Noida/Delhi)

T. Städler (2001-07; now Wissenschaftlicher Mitarbeiter, ETH Zürich)

H. Stenøien (2001; now Professor, U. of Trondheim)

S. Mousset (2002-04; Marie Curie fellow; now Maitre de Conference, U. of Lyon)

D. Presgraves (2003-04; Alexander von Humboldt fellow; now Professor, U. of Rochester)

H. Li (2003-06; now Group Leader, MPI for Computational Biology, Shanghai)

L. Camus-Kulandaivelu (2007-09; now Chargée de Recherche, CIRAD,  Montpellier)

A. Tellier (2007-12; Fellow of VW-Foundation; now Professor, TU Munich)

M. Mboup (2009-12; now DuPont de Nemours (France) SAS)

D. Živković (Fellow of VW-Foundation; now Postdoc, TU Munich)

T. Nosenko (2012-15)

A. Wollstein (Fellow of VW-Foundation; now Postdoc, LMU Munich)

B. Eldon (current)


Research Associates (Wissenschaftliche Assistenten)

E. Baake (2000-01; now Professor, U. of Bielefeld)

C. Vogl (2001-02; now Assistant Professor, Vet. med. U. of Vienna)

D. De Lorenzo (2000-06; now Lecturer, U. of Lleida, Spain)

P. Pfaffelhuber (2004-07; now Professor, U. of Freiburg)

L. Rose (2002-10; now Professor, U. of Düsseldorf)

S. Hutter (2008-15)

K. Böndel (2014-15; now Postdoc, U. of Edinburgh)